Using breed resources to improve carcass and meat quality
Paul L. Charteris and Dorian J. Garrick
Institute of Veterinary, Animal and Biomedical Sciences,
Massey University, Palmerston North
New Zealand Simmental Fielday, March, 1997
The previous year has been tough for all beef producers with low
international prices depressing returns for cattle. The catch phrases
in the beef industry of the 90's appear to be meat quality and customer
focus. Unfortunately, in New Zealand with some exceptions, few beef
producers are currently rewarded for consistently producing a high
quality beef product,. On-farm, meat quality can be improved through
changes in feeding regime such from pasture to grain finishing,
cattle management, altering selling policies and changes in breeding
policy. Breeding decisions such as breed choice, crossbreeding system
and selection of superior animals within a breed can all affect
meat quality.
1.0 Beef cattle carcass traits
Beef cattle carcass traits can be categorised as weight, yield
and meat quality related. These characteristics are of primary importance
to beef cattle farmers, processors and consumers respectively. Carcass
yield characteristics refer to the yield of saleable meat product
(meat plus specified amounts of fat) produced per carcass or per
unit carcass weight. The term meat quality can be defined in terms
of consumer requirements or more objectively such as chemical lean
content. Meat quality may include four major areas; visual quality,
eating quality, nutritional quality and safety. Some industry commentators
have described the ideal carcass as yielding a maximum percentage
of saleable meat, a minimum percentage of bone and enough fat to
meet the market requirements.
1.1 Carcass weight
With current beef cattle grading and classification, carcass weight
is the strongest determinant of price received per animal and is
therefore of great importance to the beef producer. However, inconsistent
price signals between different carcass weight ranges create difficulties
for farmers establishing target carcass weights. Unlike the farming
community, carcass weight is of lesser importance for meat processors
and of no importance to consumers.
1.2 Carcass yield
Yield traits provide an estimate of weight or percentage of saleable
beef product per carcass processed. Yield traits are important to
meat processors since an increase in saleable meat yield results
in more saleable product per carcass processed. A commonly used
measure of yield is dressing percentage (carcass weight/live weight
x 100). Since increased dressing percentage can be affected by an
increase in carcass fat or bone content, it is not a good predictor
of value of the carcass. Dressing percentage indiscriminately rewards
the deposition of lean or fat on the carcass and may encourage producers
to overfeed cattle resulting in excess fat.
Carcass muscularity is defined as the thickness of muscle relative
to skeletal dimensions and conformation is a visual assessment of
the thickness of fat and muscle in relation to skeletal measures.
Carcass conformation is an is an indirect indicator of carcass meat
yield. In New Zealand, payment per carcass is determined by conformation
rather than yield of saleable meat. A more accurate prediction of
saleable meat yield may be obtained through equations incorporating
fat depth and eye muscle area taken at the 12/13th rib intersection
in addition to carcass weight.
Subcutaneous fat depth is measured as the depth of subcutaneous
fat over the fourth quarter of the eye muscle at the 12th rib. In
practice, company graders use subcutaneous fat depth in addition
to visual assessment of fat content of the whole carcass as a guide
when determining fat class of a carcass. Measurement of the degree
of fat cover for carcass grading and selection purposes is important
since:
- Yield of lean meat decreases as fat content increases
- Consumer trends are away from high fat content meats due to
health concerns
- Fat is a waste product when it is trimmed from meat prior to
cooking.
- Consumers believe they are not receiving value for money when
a large portion of the beef product is wasted due to trimming.
- Inefficiencies arise from producing excess fat. Cost of waste
fat to the United States beef industry was estimated at US $4.4
billion dollars in 1991, $2.0 billion to produce and another $2.4
billion to transport and trim.
1.3 Meat quality
Meat quality traits are important to consumers since they affect
purchase (and repurchase) decisions and satisfaction of consumption.
Consumers perceive that yellow fat may be associated with older
animals or animals that are diseased. The chemical constituent of
pasture known to cause yellowing of fat colour have been identified
as carotenoid pigments, the primary pigment causing yellowing of
fat colour being -carotene. Cattle finished on grain consume less
carotene and so produce a non-pigmented white fat. Similarly, meat
colour is an important visual trait affecting consumer purchase
of a beef product which can be improved through feedlot finishing.
Beef that is not a bright, attractive red colour can be perceived
by consumers as not fresh, prone to spoilage or from an older animal
and therefore less tender.
Marbling is a term used to describe intramuscular fat content.
An increase in marbling is generally associated with enhanced cooking
and palatability attributes. Although there is a positive association
between marbling and beef tenderness, juiciness and flavour, the
association is weak. Recent US research has found that high levels
of marbling required to grade USDA choice were not justified in
terms of eating quality (juiciness, tenderness) of beef. In the
important beef export market of Japan, the method of cooking (broiling)
requires that beef has a high marbling content to avoid rapid cooking
and the production of offensive odours.
Consumers consider tenderness to be the most important aspect of
meat eating quality. The United States beef industry task force
identified increasing consistency and quality of beef product as
important to improving market share, with a reduction in consumer
satisfaction (primarily related to toughness) by 50% required by
1997. Marbling and connective tissue content accounted for only
20% of observed variation in tenderness, the remainder being influenced
by processes occurring postmortem, including chilling, ageing and
cooking procedures.
Other meat quality traits include flavour and juiciness. Flavour
of meat is highly influenced by environmental factors such as nutritional
regime, age at slaughter and post-slaughter management. Less desirable
flavour of beef from pasture finished as opposed to feedlot-finished
cattle has been attributed to lower fat content, or different fatty
acid composition of beef from the former group of animals. Assessment
of juiciness and flavour rely on cooking meat for a specified period
in a precise manner and subsequent sensory evaluation by a trained
taste panel.
2.0 Sources of variation in carcass and meat quality
Within any group of animals we will find variability, take a mob
of calves for example, within this mob, some calves will be heavier
at weaning due the genes they possess for growth, availability and
quality of feed they consume (both pasture and milk supply from
their dam) and their health. Now compare this mob of calves with
all the beef calves weaned in New Zealand, differences in weaning
weight are now due to genes in addition to differences in management
between farms and between regions. Thus diffences between animals
can be summarised as arising due to differences due to
- genes
- environment (nutrition, + animals health + climate + management)
When we invest in a breeding bull we are interested in his genes
he transmits to his progeny. Thus for improving carcass and meat
traits we would like to know what proportion of the total variation
in these traits are due to genetic difference between groups of
individuals. This proportion is termed heritability. Traits with
a higher heritability have a higher proportion of variability accounted
for by genes.
Significant genetic variation exists both within and between breeds,
the magnitude of this variation is generally comparable within and
between breeds for most traits. Thus, significant genetic change
can result from both within and between-breed selection. A United
sates study (Green, 1991) showed that variation within breeds for
marbling and saleable meat yield (%) was of the same magnitude as
variation between breeds for the same trait. For saleable meat yield
(kg), variation between breeds was greater than variation within
individual breeds.
Differences between breeds are more easily exploited than differences
within a breed since variation between breeds is often more highly
heritable. Say for example you wished to improve saleable meat yield
for your Hereford herd, the quickest way to accomplish this would
be to use bulls of breeds known superior for saleable meat yield
such as Limousin, Charolais and Simmental rather than select for
saleable meat yield from within the Hereford breed. In addition,
selection within breeds for such traits is often complicated by
the fact that few animals have Estimated Breeding Values (EBVs)
for traits such as saleable meat yield, marbling, meat colour and
fat colour.
Breed differences in performance characteristics are an important
genetic resource for improving efficiency of beef production and
meeting market requirements. However, more time is required before
definitive statements can be made regarding breed superiority for
a number of meat quality traits. Research comparing breeds in the
United States is summarised in Table One for which some breeds grouped
into types on the basis of four criteria.
Table One Breeds grouped into biological types for four criteria
(Cundiff et al. 1993).
| Breed group |
Growth rate and mature size
|
Lean to fat ratio |
Age at puberty |
Milk production |
| Jersey |
X |
X |
X |
XXXXX |
| Herf & Ang |
XXX |
XX |
XXX |
XX |
| Shorthorn |
XXX |
XX |
XXX |
XXX |
| Sth. Devon |
XXX |
XXX |
XX |
XXX |
| Gelbvieh |
XXXX |
XXXX |
XX |
XXXX |
| Holstein |
XXXX |
XXXX |
XX |
XXXXX |
| Simmental |
XXXXX |
XXXX |
XXX |
XXXX |
| Salers |
XXXXX |
XXXX |
XXX |
XXX |
| Limousin |
XXX |
XXXXX |
XXXX |
X |
| Charolais |
XXXX |
XXXXX |
XXXX |
X |
Increasing number of X's indicate relatively higher values, thus
Jersey cattle tend to be slower growing, have smaller mature size,
are less lean, are younger at puberty and have higher milk production
than most beef breeds.
Unfortunately there are often genetic antagonisms between traits,
this means that improvement in one trait can result in decreased
performance in another trait. For example there is a negative genetic
relationship between marbling and lean meat yield, selecting cattle
for superior marbling will tend to decrease the yield of lean meat
and vice versa. Average marbling scores and lean meat yield (%)
across breeds is shown in Figure Two. The breed with the highest
average marbling score (Jersey) also had the lowest lean meat yield,
similarly for the Chianina which had the highest lean meat yield.
These trade-offs confirm the statement that no one breed excels
for all traits it is for this reason that we have different beef
cattle breeds and utilise their genes in crossbreeding programmes.
If Chianina breeders decided to select for marbling, eight generations
(40 years) of single trait selection for this trait would be required
to increase marbling in Chianina to the level of that in Angus.
In the process however, due to negative genetic correlations, lean
meat yield of the Chianina would decrease. Due to difficulties of
measurement of marbling on the live animal, most rapid genetic improvement
would occur through crossbreeding programmes where breeds with marbling
superiority such as Wagyu or Jersey are utilised. An example crossbreeding
programme would be to mate an early maturing dam breed known for
high marbling and meat palatability to a high growth rate terminal
sire breed which produces a high lean meat yield.
New Zealand Beef Processor data
Earlier maturing breeds have a higher fat depth and lower lean
meat yield at a constant carcass weight than do late maturing breeds.
In a comparison between beef cattle breeds slaughtered at Manawatu
Beef Packers in New Zealand, Garrick (1994) found that British breeds
had higher subcutaneous fat depth at a similar carcass weight than
did Exotic x British breeds and cattle of dairy origin. The implications
of this for farmers are that steers of Continental x British breeds
can be finished to higher carcass weights without incurring fat
depth penalties compared with steers of British or Dairy x Beef
breeds.
New Zealand breed comparisons
Among the most comprehensive beef cattle breed evaluation in the
world have been conducted in New Zealand, through AgResearch, Ruakura.
In one study, 1908 steers from eleven breeds were evaluated over
five years. Sires were mated to either Angus or Hereford dams. These
trials were conducted through the 1970's, due to breed genetic trends
the relative rankings of these breeds are likely to be different
in 1997. The effect of breed of sires on carcass traits of progeny
slaughtered at 31 months age and adjusted to a fixed carcass weight
of 277 kg or a fat depth of 6.5 mm is shown in Table Two. Sire breeds
are ranked on the average yearling weight of their progeny with
highest yearling weights shown at the top of the Table.
Table Two: Effect of breed of sire on carcass traits at 31 months
of age when adjusted to a fixed carcass weight of 277 kg or a fat
depth of 6.5 mm (Morris et al. 1990)
| Sire Breed |
Fat depth (mm) |
Eye muscle area (mm) |
Hot carcass weight (mm) |
Relative carcass productivity1
|
| Maine Anjou |
4.9 |
100 |
295.8 |
103 |
| Simmental |
|
|
|
|
| German |
5.4 |
108 |
287.5 |
106 |
| French |
4.4 |
96 |
279.2 |
107 |
| Austrian |
6.2 |
101 |
276.0 |
98 |
| Swiss |
6.5 |
98 |
272.7 |
102 |
| Friesian |
6.8 |
91 |
288.5 |
107 |
| Charolais |
5.0 |
104 |
291.6 |
99 |
| South Devon |
7.1 |
96 |
282.5 |
104 |
| Chianina |
6.1 |
99 |
277.1 |
103 |
| Blonde d' Aquitane |
5.0 |
101 |
290.6 |
106 |
| Limousin |
5.4 |
104 |
273.8 |
101 |
| Hereford |
10.2 |
93 |
260.9 |
100 |
| Jersey |
8.8 |
92 |
249.2 |
94 |
| Angus |
8.4 |
97 |
249.9 |
92 |
Carcass productivity (average carcass weight x calf survival from
birth to weaning) relative to a base of 100 for Hereford x Angus
cattle. Carcass weight is hot carcass weight at 20 months of age.
A useful exercise is to highlight the five sire breeds with the
lowest average fat depth, highest average eye muscle area and highest
average carcass weight. Compare the breeds you have highlighted
in this Table with breeds which are superior for lean to fat ratio
in Table One, the two lists should match quite closely.
Summary
Carcass and meat quality traits differ in importance depending
on the sector of the beef industry examined and the requirements
of different markets. Variation in traits arises from variability
in due to both environment and genes the animals possess, however
only these genetic differences can be exploited to change meat quality
of animals in future generations. Cattle breeders can exploit genetic
variation both within and between breeds to improve carcass and
meat quality traits. However, these traits are only part of the
a large complex of traits which, together with fertility, survival
and growth performance all impact on farm profitability.
Acknowledgments
Thanks to Mr. Doug Lyneham, AFFCO, New Zealand for supplying data
from Manawatu Beef Packers. Beef cattle breeding research and extension
support is provided by the New Zealand Meat Research and development
Council (MRDC).
References
Charteris, P.L. June, 1996. Breeding Matters - Beef Supplement.
The New Zealand Animal Breeding Trust.
Cundiff, L.V.; Szabo, F.; Gregory, K.E.; Koch, R.M.; Dikeman, M.E.;
Crouse, J.D. 1993. Breed comparisons in the germplasm evaluation
programme at MARC. Proceedings of the Beef Improvement federation
Research Symposium and Annual Meeting, Asheville, NC.
Garrick, D.J. 1994. Meeting market specifications: Exploiting breeds
and crossbreeding for profitable production of beef. Proceedings
of the Central districts Sheep and Beef farmers Conference, vol
3: 35-40.
Morris, C.A.; Baker, R.L.; Carter, A.H.; Hickey, S.M. 1990. Evaluation
of eleven cattle breeds for crossbred beef production,: carcass
data from males slaughtered at two ages. Animal Production, 50:
79-92.
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